Yasako and Isako, on the run from Megamass and a powerful new searchmaton, take refuge in Megabaa's store. There, it is discovered that Densuke has the potential to open a stable portal to "the other side", now known as "Coil Domain". It is revealed that Coil Domain, Michiko, Imago and Kirabugs are remnants of an experiment gone awry, covered up by the Megamass conglomerate.